Okay, you dragged me back in one more time.
While searching for pictures of byssus openings, I came across some additional info in The Reef Aquarium Vol. 1:
Too bad we can't see the incurrent aperture very well in either picture. Squamosa clams have "numerous, large, branched tentacles" that are easily distinguished from the "small, fine tentacles" of maxima clams. That's a sure distinguishing characteristic.
Maxima clams are typically "asymmetrical and elongated, 3x longer than broad." The pictured clam is more symmetrical, which is a characteristic of squamosa.
According to RA1, squamosa clams can be confused with maxima when small because they both can have scutes on their shells. "However, those of squamosa are much larger than those of maxima and are not as closely spaced." The scutes and spacing both indicate squamosa.
The byssus opening can be that of a juvenile squamosa. Actually, it appears more like a crocea from the bottom than either squamosa or maxima. The answer might be that it's a hybrid. Squamosa is "known to form hybrids with crocea and maxima."
Nathan: If you have facts to prove me wrong, please chime in. I welcome the education.
Greg